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Z
A 0
x S ¼
x S ð z Þ ds ð z Þ
0
where s(z) is the function of distribution of the root system along the vertical cross-
section of the soil layer; A 0 is a maximum capacity of the root layer of the soil.
For conifers, r a r st ;
and along the height of the tree the value of photosyn-
thesis per unit mass of pine-needles changes negligibly, and therefore we shall
consider some point, averaged over the tree height, for which the intensity of the
photosynthetically active radiation (PAR) is determined by the Beer-Lambert law:
5K e hS L H 1
E ¼ j E E e exp 0
:
where E e is the intensity of the incoming solar radiation over the forest,
ʺ E is the
share of PAR in the total solar radiation; K e is the coef
cient of extinction.
Assuming that the temperature of pine-needles coincides with that of the
atmosphere (T L =T), we use the following approximation for the
˃
(T) function:
(
"
#
)
l
T T 1
l 1 T T 1
T 0 T 1
T 0 T 1 exp l 1
T Þ ¼max 0
;
;
where the parameter l = 1.8; T 1 is a minimum temperature at which the photo-
synthesis still takes place; T 0 is a temperature optimal for photosynthesis.
The
function characterizes the dependence of photosynthesis on soil moisture.
With a short-term over-moistening of soil the photosynthesis does not decrease, and
therefore
ʩ
Xðx S Þ ¼1
8x S 2 ½0
;
1 :
In a general case, we use the dependence:
1
;
x S 0
; x HB Þ;
Xðx S Þ ¼
ð 1 x S Þ=ð 1 x HB Þ;
x S 2 ½ x HB ;
1
˄ B ) is introduced in order to describe the strategy of distribution
of assimilates, where
ʸ i (
The function
˄ B is the biological time and to determine the ratio of the sum
of ef
cient temperatures T l to the mean multi-year sum of ef
cient temperatures T ʣ
accumulated during the vegetation period:
s B ð j Þ ¼ X
j
T l =
T R ;
l¼1
where j is the day number.
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