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measure based on the arithmetic differences between the two
matrices, D A , is given by:
K - 1
å
K
å
( FM ij ( X ) - FM ij ( M )) 2
D A ( X , M ) =
i = 1
j = i + 1
This measure is simple to calculate and does not depend on any cen-
tering protocol. However, it does not take into account the covariance
between the landmarks.
3.
Dissimilarity measure based on the Euclidean Distance
Matrix representation using relative differences: A dis-
similarity measure based on the relative differences between
matrices representing the mean form and an individual is
given by:
2
K - 1
å
K
å
æ
FM ij ( X )
FM ij ( M ) - 1
ö
ç
÷
D R ( X , M ) =
è
ø
i = 1
j = i + 1
This measure is also simple to calculate but weighs the differences dif-
ferently; e.g., the smaller distances receive more weight in the
calculation. This may or may not be desirable depending upon the
research problem.
4.
Dissimilarity measure based on logarithmic differences:
The following dissimilarity measure is based on the differ-
ences between the logarithms of the distances:
K
1
K
D
(
X M
,
)
(log
FM
(
X
)
log
FM
(
M
))
2
LR
ij
ij
i
1
ji
1
= +
5.
Dissimilarity measure based on the Euclidean
Distance Matrix representation of relative differences
in growth pattern: We have found that there are situations
in which a growth pattern can be useful in distinguishing
between groups (Richtsmeier and Lele, 1993; Richtsmeier et
al. 1993a, 1993b). There are situations in which organisms
achieve similar adult forms through differing growth pat-
terns. In this case a dissimilarity measure based on the
growth matrices (and the differences between them) can be
used. Growth data are required for each individual in ques-
tion and for each of the defined groups. The dissimilarity
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