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(Williamson & Schneider, 1993). In Proteaceae , receptacular nectaries are
common (although they may be absent in some taxa) in an alternitepalous
position between the androecium and the gynoecium (Fahn, 1979; Douglas
& Tucker, 1996) and can be considered intrastaminal. Douglas and Tucker
(1996), after developmental studies, judged these nectaries as secondary or-
gans, not reduced homologues from “lost” petal or stamen series.
Buxaceae. All members of this family have unisexual flowers. In female
flowers of some Buxus species, interstylar nectaries occur on the gynoecium
(von Balthazar & Endress, 2002). Daumann (1974) interpreted these nec-
taries as remnants of stamens, whereas Smets (1988) regards them as
convergent homologues to septal nectaries in monocots. Sarcococca and
Pachysandra lack interstylar nectaries in female flowers, but male flowers
have nectariferous pistillodes (Vogel, 1998c; von Balthazar & Endress, 2002).
Styloceras has no nectariferous tissues (von Balthazar & Endress, 2002).
Trochodendraceae. Conspicuous dorsal carpellary bulges are differentiated
as nectaries (Endress, 1986), similar to the interstylar nectaries reported for
Buxaceae.
3.8.8
Core Eudicots
In the Core Eudicots, receptacular nectaries—either continuous or fragmen-
ted—are common, located principally between androecium and gynoecium
and in association with the filament bases. Sepal, petal, and gynoecial nectar-
ies are less frequent.
Gunnerales. Flowers are wind-pollinated, reduced, and devoid of nectaries
(Wanntorp & Decraene, 2005).
Aextoxicaceae. Cronquist (1981) pointed out small receptacular nectary
glands alternating with the stamens in an antesepalous position, probably of
staminodial origin.
Berberidopsidaceae. The receptacle extends into a lobed ring-like nectary
between the androecium and the inner tepals (Decraene, 2004), and could be
described as extrastaminal.
Dilleniaceae. No nectar has been observed; flowers have pollen and are
buzz-pollinated (Tucker & Bernhardt, 2000).
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