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successful prey capture in these habitats (Daeschler
et al. 2006).
The hypotheses outlined above need to be tested
further. As the study of tetrapod terrestriality pro-
ceeds, it will be necessary to develop palaeoecologi-
cal models based on multiple lines of evidence
including sedimentology, taphonomy, functional
morphology, developmental biology, biogeochem-
istry and other disciplines that can be synthesized
to provide a holistic picture of these ancient non-
analogue ecosystems.
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lycophyte Haskinsia Grierson and Banks from
the Devonian of western Venezuela, with obser-
vations on leaf morphology and fertile specimens.
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aggrading sand-bed braided Niobrara River, Nebraska:
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