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d 3
P 4
P 1
P 3
d 1
d 2
P 2
45°
SX
45°
FIGURE 6.4
Wandering quarter sampling (Catana, 1963). S is a randomly selected point. P 1 is the near-
est object to S within a quarter determined by the direction of the transect (indicated by the
arrow), having S as the starting point for searching. The nearest object to P 1 (i.e., P 2 ) found in
the next quarter determines the first measured distance d 1 used in the formula for density
estimation. The procedure continues in a similar fashion, producing additional distances d 2 ,
d 3 , . . . . See text for further details.
be the mean distance between items. If items in the study region are ran-
domly dispersed, the density is estimated as
ˆ
DAd
/
2
,
(6.6)
wq
where A is the unit of area, and d 2 is interpreted as the mean area of all
items. For a clumped spatial distribution of items, Catana (1963) developed
an estimator of D wq based on methods suggested by Cottam et al. (1953, 1957),
that is,
ˆ wq
DNN
itc ,
cl
where N cl is the number of clumps per unit area, and N itc is the number of
items per clump. However, here only the estimator of D wq for randomly dis-
persed items is considered.
As noted by Diggle (2003), the wandering-quarter method is an ingenious
method whose slow adoption may be because of the use of longer chains of
object-to-object distances that would produce distances with increasing like-
lihood of boundary problems (i.e., reaching the end of the region in less than
n steps either because Catana recommended n = 25 or because of encounter-
ing the side boundary). Another reason why Catana's method is not so popu-
lar is the lack of an easy-to-compute standard error of the estimate. Hall et al .
(2001) proposed a bootstrapping procedure for a generalized version of the
wandering-quarter method, but the density estimator is different from that
suggested by Catana (1963).
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