Agriculture Reference
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GES
O
O
OH
OH
O
O
Geranyl acetate
Geraniol
Geranial
Geranic acid
Neral
O
OOH
Neric acid
O
OH
O
Citronellol
Nerol
Citronellyl acetate
OH
O
O
O
OH
Rose oxide
Citronellal
Citronellic acid
(b)
(c)
1.2
Control
1.0
0.8
0.6
0.4
0.2
0.0
Transgenic
Control
Transgenic
(d)
(e)
(f)
12%
9%
16%
35%
16%
60%
68%
5%
79%
Smell
Retronasal
aroma
Preference
Plate 5.
Metabolic engineering to improve tomato fruit aroma quality (Davidovich-Rikanati, R., Sitrit, Y., Tadmor,
Y., Iijima, Y., Bilenko, N., Bar, E., Carmona, B., Fallik, E., Dudai, N., Simon, J.E., Pichersky, E. and Lewinsohn, E.
(2007) Enrichment of tomato flavor by diversion of the early plastidial terpenoid pathway.
Nature Biotechnology
25(8), 899-901). (a) The metabolism of geraniol in geraniol synthase (GES)-expressing tomato. (b) Phenotypes of
controls and transgenic tomato fruit. (c) Contribution of geraniol derivatives to the aroma of transgenic tomato.
(d-f) Organoleptic evaluations of transgenic tomato by untrained panellists.
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