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data indicate that sexual reproduction may at least sometimes play a significant
role in the propagation of this orchid.
Reproductive success of T. trianthophora may be limited by predation. Keenan
( 1996 ) reported that an unspecified number of flowers and developing capsules
were routinely lost to herbivores in New Hampshire. Willems ( 1994 ) found chip-
munks and most particularly slugs to be serious predators of young plants in
Massachusetts. Rapid seed maturation, usually occurring within 7 days after fertil-
ization (Pridgeon et al. 2005 ), may reduce the loss to predators. In addition, devel-
opment, logging, and other alterations of the habitat pose a threat. The small size of
the plants and many of the populations also render them susceptible to stochastic
events and inadvertent disturbances (Ramstetter 2001 ).
References
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Hampshire, Durham, NH
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Proceedings of the 11th World Orchid Conference. The American Orchid Society, Miami,
pp 121-135
Dressler RL (1993) Phylogeny and classification of the Orchid Family. Dioscorides, Portland, OR
&REUDENSTEIN *6 'YNOSTEMIUM STRUCTURE AND RELATIONSHIPS OF THE #ORALLORHIZINAE
(Orchidaceae: Epidendroideae). Plant Syst Evol 193:1-19
Keenan PE (1986) New stations for Platanthera flava and Triphora trianthophora and other obser-
vations. Rhodora 88:409-412
Keenan PE (1988) Three-birds orchids at Golden Pond. Am Orchid Soc Bull 57:24-27
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Keenan PE (1996) Further observations on Triphora trianthophora - Part 4. N Am Nat Orchid
J 2:196-207
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Luer CA (1975) The native orchids of the United States and Canada excluding Florida. New York
Botanical Garden, New York
Medley ME (1979) Some aspects of the life history of Triphora trianthophora (Sw.) Rydb. (three
birds orchid) with special reference to its pollination. M. S. thesis, Andrews University, Berrien
Springs, MI
Medley ME (2002) Triphora Nuttall. In: Flora of North America Editorial Committee (ed) Flora
of North America North of Mexico, vol 26. Oxford University Press, New York; Oxford,
pp 592-595
Porcher RD (1977) The rediscovery of Triphora trianthophora , three birds orchid, in the Coastal
Plain of South Carolina. Castanea 42:108-111
Pridgeon AM, Cribb PM, Chase MW, Rasmussen FN (2005) Genera Orchidaceum, vol 4.
Epidendroideae (part 1). Oxford University Press, New York
Ramstetter JM (2001) Triphora trianthophora (Swartz) Rydb. (Three-birds orchid). Conservation
and Research Plan. New England Wild Flower Society, Framingham, MA
Sheviak CJ (1974) An introduction to the ecology of the Illinois orchidaceae. Illinois State Museum
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Willems SA (1994) Observations on reproduction in Triphora trianthophora (Orchidaceae).
Rhodora 96:30-43
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