Biomedical Engineering Reference
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(b)
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FIGURE 1.17 Hassenstein-Reichardt model of motion processing: (a) preferred direction, (b) null direction, and (c) motion
director. Adapted from Refs. 55-57 .
the view as seen by photoreceptor A is time
delayed by τ and then multiplied by the immedi-
ate response from photoreceptor B. If the two
photoreceptors have seen the same stimulus,
there will be a high correlation, as indicated by
a large response from the multiplier. The left-to-
right motion is said to be in the preferred direction
for this arrangement see Figure 1.17 a. On the
other hand, a stimulus traveling from right to
left (termed the null direction ) will have a mini-
mal response from the multiplier stage; see
Figure 1.17 b.
To detect motion in both directions for a given
pair of photoreceptors, a mirror-image configu-
ration of hardware is added; see Figure 1.17 c.
This results in a large positive response in the
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