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A
B
O 2
Shelf-sea
chalk
Th
common
Deep-sea
chalks
Th rare or
absent
Ch
Zo
Te
Pl
Th
FIGURE 3 Schematic representation of typical ichnofabrics and recurring trace-fossil suites repre-
senting the Zoophycos Ichnofacies in chalks deposited in (A) well-oxygenated deep-sea versus
shallow-shelf settings, and (B) variably oxygenated basins (Ch; Chondrites ,Zo; Zoophycos ,Te;
Teichichnus ,Pl; Planolites ,Th; Thalassinoides ). As shown in bothA and B, the expression of the trace
fossils is improved in heterogeneous successions at transitions between lighter and darker beds (bed-
junctionpreservation). As depicted inB, systematic changes in trace-fossil diversity, burrowdiameters,
and depths of burrow penetration with changes in benthic oxygenation result in the stacking of distinct
oxygen-related ichnocoenoses. Note in B that burrows representing one ichnocoenosis may overprint
laminae or ichnofabrics formed under previous conditions. Examples of deep-sea, shelf, and oxygen-
deficient epeiric basin chalk ichnofabrics are provided in Figs. 4, 5 and 6 , respectively.
Tiering is particularly well manifested by cross-cutting relationships among
recurring ichnotaxa in outer-shelf and epeiric basin chalks (e.g., Ekdale and
Bromley, 1982, 1991; Frey and Bromley, 1985; Locklair and Savrda, 1998a,b ;
Fig. 3 ). Thalassinoides typically is overprinted by all other ichnotaxa and, thus,
characterizes a shallow tier. In contrast, Chondrites is superimposed on all other
structures (e.g., Fig. 5 C), reflecting its prevalence in the deepest tier. Other recur-
ring ichnotaxa ( Phycosiphon , Planolites , Taenidium , Teichichnus , and Zoophy-
cos ) represent intermediate tiers.
The substrate composition and consistency may have an impact on ichno-
coenoses and ichnofabrics. As carbonate contents decrease substantially, either
due to increased influx of siliciclastic muds or enhanced carbonate dissolution
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