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FIGURE 2 Common deltaic trace fossils. (A-C) Tidal-deltaic facies, Middle Jurassic Lajas For-
mation, Neuqu ยด n Basin, Argentina. (A) Ophiomorpha (Oph) in delta-front facies. (B) Taenidium
(Ta). (C) Thalassinoides (Th) at a parasequence boundary. (D-F) Wave-influenced facies, Creta-
ceous Ben Nevis Formation, offshore Newfoundland, Canada. (D) Ophiomorpha (Oph). (E) Ophio-
morpha (Oph), Skolithos (Sko), and Teichichnus (Te). (F) Asterosoma (Ast) and Ophiomorpha
(Oph). (G-H) Gilbert-type delta, Pleistocene, Conway Flats, North Canterbury, New Zealand.
(G) Diplocraterion (Diplo). (H) Burrow mottling, Planolites (Pl) and Phycosiphon (Phy). (I) Tha-
lassinoides (Th) in channel facies, Cretaceous Blackhawk Formation, Book Cliffs, USA.
(3) the ability for one organism to create multiple burrow morphologies (behav-
ioral plasticity) and to create extensive burrow networks (e.g., Thalassinoides );
and (4) changes in ichnodiversity through geologic time (e.g., colonization of
brackish-water environments, Buatois et al., 2005 ). While ichnodiversity
cannot be used as a direct indicator of species richness, it is a valid proxy for
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