Fluvial Environments - Trace Fossils as Indicators of Sedimentary Environments - page 338

FIGURE 6 Trace fossils that can be assigned to the Grallator (AandB), Batrachichnus (CandE), and
Brontopodus Ichnofacies (D). (A) Abundant bird-like tracks (B3-4 sl) of the ichnogenus Gruipeda pre-
served as positive hyporeliefs. (B) Bipedal trackway composed of large tridactyl footprints (B 3-4 tk,
arrows), preserved as positive hyporeliefs. (C) Manus-pes set of the ichnogenus Batrachichnus (Q4/5),
negative epirelief. (D) Several Ichniotherium footprints (Q 5/5 tk) in a mud-cracked sandstone sole
(positive hyporelief). (E) Manus-pes set of the ichnogenus Brachychirotherium (chirotheriid tracks),
preserved as positive hyporelief. Photographs are from the Early Jurassic Santo Domingo Formation,
La Rioja province, Argentina (A); from the Oligocene Vinchina Formation in La Rioja province,
Argentina (B); from the Permian Carapacha Formation in La Pampa province, Argentina (C); from
the Permian Tambach Sandstone, Germany (D); and from the Middle Triassic Tarjados Formation,
La Rioja province, Argentina (E). m, manus; p, pes.

trackways). The Batrachichnus Ichnofacies, as represented in fluvial deposits
( Fig. 1 H), is dominated by different quadrupedal trackways (Q 4/5, Q 5/5 h,
Q 5/5 large, or chiroteriid tracks; see Table 1 ). Associated undiagnostic trace
fossils include arthropod trackways, meniscate burrows, simple horizontal
trails, horizontal burrows, bilobed burrows/trails,

trough-like reliefs, and