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FIGURE 7 General trace-fossil distribution in glacial environments. Ar : Arenicolites ; Co :
Cochlichnus ; Ch : Chondrites ; Cn : Conichnus ; Di : Diplichnites ; Dc : Diplocraterion ; Dp :
Diplopodichnus ; Dt : Diopatrichnus ; Ga : Gastrochaenolites ; Go : Gordia ; Me : Mermia ;
Pa : Palaeophycus ; Pl : Planolites ; Rh : Rhizocorallium ; Sk : Skolithos ; Si : Siphonichnus ; Te :
Teichichnus ; Th : Thalassinoides ; Un : Undichna . After Buatois and Ma´ngano (2011) .
Table 2 in http://booksite.elsevier.com/9780444538130 for ichnotaxa and refer-
ences; Fig. 7 ). They generally show a low degree of bioturbation, impoverished
diversity, and diminutive size, when compared with ichnofaunas from normal-
saline environments. These features are typical of brackish-water ichnofaunas
(e.g., Buatois et al., 2005 ). However, the presence of truly marine, deep-water
ichnogenera (e.g., Phymatoderma , Zoophycos ) suggest the establishment of
the archetypical Cruziana Ichnofacies in deep-water settings, impoverished
due to lowered salinity owing to dilution by glacial meltwaters. The occurrence
of the Cruziana Ichnofacies in turbidites, instead of the expected Nereites Ichno-
facies, can be explained by the downslope transport of food by turbiditic currents
and mass flows ( Eyles et al., 1992 ).
4. ICHNOLOGY OF RECENT GLACIAL ENVIRONMENTS
Little ichnological research has been performed in modern glacial environ-
ments, all of them concentrated in the Arctic regions. The record of bioturbation
in glaciolacustrine and terrestrial environments includes chironomid traces from
sediments of a Norwegian lake in contact with ice ( Duck and McManus, 1984 ),
vertebrate and invertebrate traces of a fluvial pointbar in northern Alaska com-
pared to Treptichnus , Cochlichnus , Archaeonassa ( Aulichnites in Martin, 2009 ),
and Helminthoidichnites ( Martin, 2009 ), and a low-diversity Palaeophycus -
dominated trace-fossil assemblage with rare Arenicolites from postglacial
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