Environmental Engineering Reference
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FIGURE 5 Crustacean burrows and their tracemakers (inset photos). (A) Unlined Skolithos (Sk) in
sand at the intertidal/supratidal transition. Burrows are made by the amphipod, Talorchestia longi-
cornis . (B) Large-diameter Thalassinoides (Th) produced by the lobster Homarus americanus in
firm mud in 10 m water depth (photo courtesy of Tyler Hauck). (C) Diplocraterion (Di) produced
by the amphipod Corophium volutator in firm mud. (D) Corophium sp. generated Arenicolites (Ar)
in fine-grained sand. Scale bar is in centimeters. (E) Smooth-walled Thalassinoides made by
the shrimp Upogebia pugentensis in sand. (F) Smooth-walled Thalassinoides made by the shrimp
Neotrypaea californiensis in firm mud. Scale bar is in centimeters. (G) Psilonichnus (Ps, white
arrows) produced by the crab Hemigrapsus sp. in sand.
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