Environmental Engineering Reference
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A
B
30 m m
30 m m
C
D
100 m m
10 m m
E
F
30 m m
300 m m
FIGURE 7 SEM images of casted microborings produced by various chemotrophs. (A) Orthogo-
num lineare , recent, Faial, Azores, 500 m water depth. (B) Flagrichnus baiulus , Svalbard, Norway.
(C) Scolecia serrata , Carboniferous, Buckhorn Asphalt Lagerst¨tte, USA. (D) Semidendrina
pulchra , recent, Faial, Azores, 60 m water depth. (E) Entobia mikra , Pleistocene, Rhodes, Greece.
(F) Imergentia isp., Pleistocene, Rhodes, Greece.
The ichnofamily Dendrinidae bears a number of distinct fan-shaped traces, the
identity of which is not entirely solved either, even though the most likely candi-
dates are naked foraminiferans as discussed for Semidendrina pulchra ( Bromley
et al., 2007 ; Fig. 7 D). Fossil representatives of this ichnofamily comprise the
ichnogenera Dendrina , Clionolithes , Dendrorete , Nododendrina , Platydendrina ,
Ramodendrina , Pyrodendrina ,and Hyellomorpha , which are most diverse in the
Devonian and Cretaceous ( Hofmann, 1996; Vogel et al., 1987 ).
With respect to the important group of mainly macroboring sponges, two
dwarf forms have been described, as there are the ichnospecies Entobia mikra
( Fig. 7 E) and Entobia nana ( Wisshak, 2008 ), the biotaxonomical identities of
which have not been revealed as yet.
Finally, there is a diverse set of bioeroding ctenostome bryozoans that have
produced characteristic networks of
large microborings since the Early
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