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30
20
10
0
5
500
1000
PPFD ( µ mol m 2 s 1 )
1500
2000
Figure 8.2 Light response curves of mid-shoot leaves from young
'Fuji' apple trees grown under full daylight ( ),52.9% daylight
(
),34.2% daylight ( ),15.8% daylight ( ) and 5.0%
daylight (
). From Asada and Ogasawara (1998). Reproduced
with permission. Under natural light conditions 2000 µ mol m 2 s 1
PAR corresponds to c. 1000 W m 2 of total radiation measured by
radiometer or 100 Klux measured with a light meter.
complex. 'Sun' leaves in exposed (exterior) canopy positions have higher net
photosynthesis per unit light and higher stomatal conductance (Campbell et al. ,
), but also have numerous other adaptations leading to higher photosyn-
thetic potential. Humidity may control stomatal conductance directly (see
Chapter
),hencephotosynthesis,andinothercasesphotosynthesis
appears to control stomatal behaviour rather than vice versa (Lakso,
,pp.
-
).
Light response curves
Apple leaf photosynthesis is of the C type with a hyperbolic light response that
typically saturates at
mol quanta m s (Figure
). The light
compensation point, i.e. the light level below which net CO exchange is nega-
tive,withrespirationexceedingphotosynthesis,is
-
µ
.
molquantam s .
Good rates of photosynthesis per unit leaf area for healthy exposed leaves are
around
-
µ
µ
mol CO m s (Lakso,
) although much higher rates
mg CO dm h by Avery (
have been recorded, e.g. over
), about
mg CO dm h by Bravdo (
mg CO dm h by
) and
-
Looney (
).
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