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biological process, it awaits rigorous testing by the tools of molecular genetics.
The model in its present form is capable of universal computation. This hints at
future directions and the use of ciliates as model systems for exploring cellular
computation.
Acknowledgments This research was partially supported by the Natural Sciences and
Engineering Research Council of Canada grant R2824AO1 to L.K. and National Insti-
tute of General Medical Sciences grant GM59708 to L.F.L. We thank Mark Daley for
comments and Jeremy Newton-Smith with help in preparing the manuscript.
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