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intensity of swan grazing and movements of the
foraging birds. Swan grazing typically occurs from
May to September and involves several flocks
(Trump, 1996), so a more comprehensive IBM
which covers an entire year and involves larger
areas of river catchment is required if the model is
to be applied effectively. Elsewhere, the versatility
of IBMs has been demonstrated by their use in
predicting the effects of environmental change
on a range of animal populations, including
river-feeding swans, coastal wading birds and
invertebrate-feeding fish (Stillman and Goss-
Custard, 2010; West et al ., 2011). IBMs can
also be used in freshwater ecosystems to model
resource-consumer interactions. Such models
could be used in a decision-support role and
should be of particular interest to those with
wildlife, environmental and fisheries interests
involved in river conservation and management.
conservation of natural habitats and of wild fauna and
flora. Official Journal of the European Communities L206 :
7-50.
Dawson FH (1989) Ecology and management of water
plants in lowland streams. Report of
the Freshwater
Biological Association 57 : 43-60.
Environment Agency (2004) The State of England's Chalk
Rivers. A report by the UK Biodiversity Action Plan
Steering Group for Chalk Rivers , Environment Agency,
Bristol.
O'Hare MT, Stillman RA, McDonnell J, Wood LR
(2007) Effects of mute swan grazing on a keystone
macrophyte. Freshwater Biology 52 : 2463-75.
Stillman RA (2008) MORPH - an individual-based model
to predict the effect of environmental change on
foraging animal populations. Ecological Modelling 216 :
265-76.
Stillman RA, Goss-Custard JD (2010) Individual-based
ecology of coastal birds. Biological Reviews 85 : 413-
34.
Trump DPC (1996) Impact of mute swans ( Cygnus olor )
on agricultural and fishing interests - Wylye and Upper
Avon valleys, Wiltshire. MAFF commissioned R&D
project VC0105. ADAS, Bristol.
van Gils JA, Gyimesi A, van Lith B (2007) Avian
herbivory: an experiment, a field test, and an allometric
comparison with mammals. Ecology
Acknowledgements
The authors thank the Freshwater Biological
Association for access to the river. Kevin Wood was
supported by a Centre for Ecology and Hydrology
Algorithm (Natural
88 :
2926-
35.
van Gils JA, Beekman JH, Coehoorn P, Corporaal E,
Dekkers T, Klaassen M, van Kraaij R, de Leeuw R, de
Vries PP (2008) Longer guts and higher food quality
increase energy intake in migratory swans. Journal of
Animal Ecology 77 : 1234-41.
West AD, Stillman RA, Drewitt A, Frost NJ, Mander M,
Miles C, Langstone R, Sanderson WG, Willis J (2011)
WaderMORPH - a user-friendly individual-based
model to advise shorebird policy and management.
Methods in Ecology and Evolution 2 : 95-8.
Environment
Research
Council) studentship.
References
Berrie AD (1992) The chalk stream environment.
Hydrobiologia 248 :3-9.
Council of the European Communities (1992) Council
Directive 92/43/EEC of 21 May 1992 on the
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