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Fig. 2.6 Typical light-
response curves of early,
mid and late successional
species are shown. (From
Bazzaz 1979)
30
Early
20
Mid
10
Late
0
3
6
9
Light Intensity (1000 ft-c)
-10
140
slope=0.701***
120
Ad
100
Ca
Cj
80
Cs
Cs *
Ma
60
Xm
Ol
Cp
Qm
Mt
40
Ns
Dp
Qg
Qr
Tc
As
Cu
Bn
20
Pv
0
0
50
100
150
200
250
LMA (g m −2 )
Fig. 2.7 Leaf maturation period and leaf mass area across different evergreen broad-leaved tree
species (Miyazawa et al. 1998): Ad, Actinidia deliciosa ; As, Annona spragu e i ; Bn, Brassica
napus ; Ca, Coffea arabica ; Cp, Connarus panamensis ; Cs, Castanopsis sieboldii ; Cu, Cucumis
sativus ; Dp, Desmopsis panamensis ; Ma, Morisonia americana ; Ol, Ouratea lucens ; Qr, Quercus
rubra ; Tc, Theobroma cacao ; Xm, Xylopia micrantha . Open squares , species attaining full pho-
tosynthetic capacity before full leaf expansion; open triangles , species attaining full photosyn-
thetic capacity at full expansion; closed squares , delayed greening; d , days
only in August (Matsumoto 1984); in Pinus pumila , full photosynthetic capacity is
attained only in September or even the following spring (Kajimoto 1990). Evergreen
broad-leaved trees such as Machilus thunbergii , Castanopsis sieboldii , and
Quercus glauca show similar delay in foliar development (Kusumoto 1961;
Miyazawa et al. 1998). In general, broad-leaved evergreen species with heavier, longer-
lived leaves take longer to develop their full photosynthetic capacity (Miyazawa
et al. 1998; Fig. 2.7 ).
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