Herpetofauna of the eastern African forests - Biogeography and Ecology of the Rain Forests of Eastern Africa - page 185

Table 9.2. Subspecies of rep tiles ound in both the easten and westen orests of Arica

Subspecies Locaion

Holaspis guentheri lavis (Werner, 1895)

northeast Tanzania south to Amatongas and Dondo,
Mozambique

Holaspis guentheri guentheri (Gray, 1863)
Na triciteres vaiega ta sylvatica (Broadley, 1966)
Na triciteres variegata variegata (Peters, 1861)

Uganda, southwest to Angola, northwest to Sierra Leone
forested areas of southern Tanzania, south through
Mozambique and eastern Zimbabwe to Zululand
Cameroun west to Guinea

Bitis gabonica gabonica (Dumeril & Bibron, 1854)

forested areas from Nigeria east to southen Sudan,
Uganda and westen Kenya south to Angola, Zambia,
eastern Zimbabwe and Zululand

Bitis gabonica rh inoceros (Schlegel, 1855)
Atheis nitschei nitschei (Tonier, 1902)

Guinea to Ghana
western Tanzania, Rwanda, Burundi, Uganda and adjacent
Zaire

Atheris nitschei runweensis (Bogert, 1940)

north from Rungwe to Kigoma; northeastern Zambia and
northen Malawi (see Rasmussen & Howell, 1982)

able to survive relaively dry condiions, such as
those found at the forest edge, and may even
persist in some areas where forest has recently
been removed.

those which are restricted to single mountain
blocks.
Those species found along much of the length
of the Easten Arc mountains include the anurans
Athroleptis ai nis, Buo brauni, Neaophynoies
toniei, Afixalus u/uguuensis, Hp eroius punai-
ulatus, Lptopelis barbouri, Lptopelis parkeri,
Lptopelis uluguuensis, Athrolptides matiensseni
and Callulina krii, the caecilian Scolecomophus
vittatus and the snakes Tp hlops ierrai and Atheris
ceratophous. As already noted at the ime of
Loveridge, many of these were considered to be
Usambara/Uluguru endemics, but recent collect-
ing has revealed their presence in the Uzungwa
Mountains.
Species known from only the Usambara and
Uluguru and outlying mountains include the frog
Schoutedenella xenoctyla, the geckos Urocoyleon
woltestofi and Lygoacylus conradti, and the
snakes Geodipss vauerocegae and Elapsoidea nira.
The latter is also recorded from the Nguru
Mountains, a poorly collected area which may
well yield more species found in both the Usam-
baras and Ulugurus as its intermediate posiion
might suggest.
Three species are known to occur only in the
Uluguru and Uzungwa mountains: the frog
Phynobatrachus uzungwensis is at present known

Sp eies with a Ta nganyika-Nyasa ystem
distribution patten

This category may be further broken down into
those species which occur in one block of or along
the length of the Tanganyika-Nyasa system, and
those which are restricted to the forests of the
Eastern Arc.
Species which extend into the southern porion
of the Tanganika-Nyasa system to the Malawi
forests are the frogs Arthro/eptis reichei and
Hy perolius spinigularis, the caecilian Scolecomophus
kirkii, the chameleons Chamaeleo goetze and
Rhampholeon nchisiensis and the snake Crota-
phopeltis tonieri.
A single species of anuran, Arthroleptis francei,
appears to be isolated in forest on Mulanje
Mountain (Poynton & Broadley, 1985a).
Species limited to the Easten Arc mountains
of Tanzania fall into several groups: species which
occur in several of the forests, those which occur
in only the northen or southen porions, and