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TABLE 3.
q=0
f
(
T
) = 1/D
t
q=1
f
(v) =
f
1
+
f
2
-
tf
1
f
2
q=2
f
(•) =D
tf
1
f
2
q=3
f
(
C
) = 0
taking more and more cross-products of intensities into consideration until
for the strongest logical function being short of contradiction—the logical
“AND”—the single cross-product of all stimuli is computed.
We have carefully avoided associating with “
p
” or with “
f
” the actual
output frequency. We called
f
the “internal frequency”. The reason will
become obvious in a moment. It is well known that after the production of
each pulse a physiological neuron requires a certain moment of time D
t
R
—
the so-called “refractory period”—to recover from its effort and to be ready
for another pulse (see also fig. 14). But in table 3 the resulting frequency
for zero threshold is the reciprocal of the pulse duration D
t
, which, of course,
is unmanageably high for a neuron whose refractory period is clearly much
longer than the duration of its pulse
(29)
DD
t
R
>
t
.
It is very easy indeed to accommodate this difficulty in our calculations, if
we only realize that the actual output frequency
f
o
of the element is the fre-
quency
f
at which it “wants” to fire, reduced by the relative time span in
which it cannot fire:
(
)
(30)
f
=-
f
1
f
D
t
R
.
o
o
Solving for the output frequency in terms of internal frequency and refrac-
tory period we have:
f
ft
R
f
o
=
.
(31)
1
+
D
From this is easily seen that the ultimate frequency at which an element
can fire is asymptotically approached for
f
Æ 1/D
t
, and is given by:
+
1 1
f
omax
=
DD D
,
(32)
t
t
t
R
which is in perfect agreement with our concept of the physiological behav-
ior of this element. The actual values for pulse duration and refractory
period may be taken from appropriate sources (Eccles, 1952; Katz, 1959).
3.2. The Neuron as an “Integrating Element”
The element discussed in the previous paragraphs is an “All or Nothing
Device”
par excellence
, and in the two subtitles “Synchronism” and “Asyn-
chronism” we investigated only how this element behaves when subjected
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