Biology Reference
In-Depth Information
current generation are completed), if multiple offspring have dispersed
to a particular point, only one of them will randomly be chosen to occupy
that site in the next generation, the others dying. Offspring production and
dispersal are conditioned by other NEWGARDEN input conditions (e.g.,
age-specifi c pollen production, mortality).
Microgamete-Pollen Dispersal Distance Values: EasyPollen Statements
As with offspring dispersal, pollen dispersal distances are defi ned by the
user as a hierarchically nested set of frames of distances from which pollen
is received by a particular female in a given mating. When a particular
female is selected for a mating, NEWGARDEN selects a pollen donor frame
according to the probability assigned to that frame. Then, an eligible pollen
donor for that mating is chosen at random from within the frame. Notice
that offspring are dispersed to available grid points at random, while pollen
distance is determined by fi rst selecting a female for a particular mating, and
then conditioned by other microgamete-related input specifi cations (e.g.,
age-specifi c pollen production), choosing one of the eligible pollen donors
at random within the distance frame selected with the given probability.
Since these two processes are different, the input code for pollen dispersal
distance differs from that used to specify offspring dispersal distance frames.
For pollen dispersal distance, EasyPollen statements are used:
<EasyPollen>
<pollenframe low=“0” high=“10” prob=“0.35”/>
<pollenframe low=“11” high=“25” prob=“0.35”/>
<pollenframe low=“26” high“55” prob=“0.25”/>
<pollenframe low=“56” high=“701” prob=“0.05”/>
<pollenframe low=“702” high=“Inf” prob=“0.0”/>
</EasyPollen>
The fi rst pollen frame statement above describes the fi rst eligible pollen
donor frame as a square centered on the female selected for a given mating,
with four sides (right, above, left, and below) each 9 units horizontally or
vertically from the central female. Thus, the distance of a side from the
central female is the (“high=” value)—1. In other words, the fi rst pollen
frame runs from the grid point holding the female involved in the mating,
out to 9 grid points in the positive or negative x and y directions. The
probability statement (prob=“0.35”) designates a 35% chance that the
pollen donor will be selected from within this frame. The fourth pollen
frame is bounded by a smallest square with sides at 55 units, and the largest
square of the frame at 700 units, in positive or negative directions from the
female. There is a 5% chance that a pollen donor will be chosen at random
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