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Table 6.20 Trophic pyramid of the Okhotsk Sea ecosystem considered in calculations of
biocomplexity indicator
Consumers of energy and
matter
Sources of energy and matter
B
1
B
2
B
3
B
4
B
5
B
6
B
7
B
8
B
9
B
10
B
11
B
12
B
13
B
14
B
15
B
16
B
17
B
18
B
19
Phytoplankton, B
1
0000000000000000010
Bacterioplankton, B
2
0000000000000000001
Microzoa, B
3
1100000000000000001
Microzoa, B
4
1100000000000000000
Carnivores, B
5
0111100000000000001
Zoobenthic animals, B
6
1111100000000000001
Flat-
sh, B
7
0000011111111111100
Cof
dae, B
8
0000111111111111100
Ammodytes hexapterus, B
9
0000011001110001000
Mallotus, B
10
0000111000100000000
Theragra chalcogramma, B
11
0000100001110000000
Salmonidae, B
12
0000011001111001000
Coryphaenoides, B
13
0000111111111111100
B
14
0000111111111111100
B
15
0000110000000010000
Crabs, B
16
0000010111010001100
Laemonema longipes, B
17
0000110000000000100
Biogenic salts, B
18
0000000000000000001
Detritus, B
19
1111111111111111100
People, B
20
1111111111111111111
Notations B
14
Reinchardti ushippoglossoi des matsuurae, B
15
Clupeapallasi pallasi Val
component at the expense of the m-th component biomass; k
im
(i = 18, 19) is the
transformation coefficient from the m-th coponent to the i-th component; ki20
i20
is the
characteristic of anthropogenic in
fl
uence on the i-th component; Si
i
={i: x
ij
=1,
j =1,
, 19} is the food spectrum of the i-th component, T
W
is water temperature;
S
W
is water salinity; B
m,*
= max{0, B
m
−
B
m,min
};
…
1
;
if B
m
B
m
;
min
x
ij
¼
0
;
if B
m
\
B
m
;
min
Designate the water area of the Okhotsk Sea by
Ω
={(
φ
,
ʻ
)}. The value of the
biocomplexity indicator for any area
ʾ
ˉ
(z
1
, z
2
, t), where [z
1
,
z
2
] is the water layer located between depths z
1
and z
2
. The maximal value of
ʾ
ˉ ∈ Ω
is determined by
20) is reached during the spring-summer period, when nutritive relations
into the Okhotsk Sea ecosystem are extended, the intensity of energetic exchanges
is increased and horizontal and vertical migration processes are stimulated. In the
winter time the value of
=
ʾ
max
(
≈
ʾ
is reduced to near
ʾ
min
(
≈
8). The spatial distribution of
re
fl
ects the local variability of the food spectrum for its components. Table
6.21
and
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