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(b)
Figure 3.2.
(cont.)
the most SO 2 tolerant in Britain (Hawksworth & Rose 1970 ; Laundon 1973 ;
Seaward & Hitch 1982 ). L. conizaeoides prefers acidic bark (Tønsberg 1992 ; Wirth
1999 ; Massara 2004 ). It was formerly widespread in lowland areas of Europe
subjected to air pollution and reported from several eastern North American
cities and near sulphur springs in Iceland (Purvis et al. 1992 ). Its occurrence
on birch (Betula) in Iceland near sulphur springs has been called into question
and is believed to reflect an unintentional introduction (Bailey 1968 ; Laundon
2003 ). Laundon ( 2003 ) considered the lichen to be exceptional for being
unknown before 1860. He studied many eighteenth and early nineteenth
century herbaria to locate early specimens without success. The earliest speci-
mens he recorded came from two disjunct areas. L. conizaeoides was originally
collected in England c. 1862 at Twycross, Leicestershire, by the Reverend
A. Bloxam (BM) at a time when the county was probably beginning to experience
the effects of background air pollution (Laundon 2003 ). A second collection
was made in 1865 from larch (Larix) from Mittersill in the Salzach valley in
the Austrian Alps (Laundon 2003 ). L. conizaeoides is probably native to southern
Central Europe where it grows in woods with dwarf mountain pine (Pinus mugo),
especially in boggy areas (Wirth 1985 ). The species is believed to have reached
England by windborne propagules and was widespread by 1880 (Laundon 2003 ).
L. conizaeoides may have colonised the railway network first ( Fig. 3.3B ) , espe-
cially track-side fences along the smoky, windy corridors created by steam
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