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100.000
(a) Postfire chaparral
Ec = 7.67
(b) Postfire chaparral
Ec = 10.82
10.000
1.000
0.100
0.010
0.001
100.000
(c) Mature chaparral
Ec = 2.11
(d) Mature chaparral
Ec = 1.73
10.000
1.000
0.100
0.010
0.001
100.000
(e) Australian heath
Ec = 42.59
(f) Australian heath4
Ec = 47.26
10.000
1.000
0.100
0.010
0.001
0 1020304050607080
Rank sequence
0 1020304050607080
Rank sequence
Fig. 11.4 Dominance-diversity curves for (a,b) Californian and (c,d,e,f) Australian shrublands
based on cover (from Keeley & Fotheringham 2003b ). Californian sites fit a geometric model
and Australian sites showed a significant fit to MacArthur's broken stick model. Ec, Whittaker's
equitability index.
species, being found in a very small fraction of quadrats within a site as well as
within a small fraction of sites (Keeley et al. 2005c ). Based on dispersal capacity
this does not appear to be driven by metapopulation dynamics but rather micro-
habitat specialization.
In contrast, Australian heathland species are more equitably distributed, as
illustrated by their narrow species-abundance distribution and dominance-
diversity relationship that fits MacArthur's broken stick model ( Fig. 11.4c , d ).
The implication of this model is that populations reach a stable equilibrium
without the development of dominance by any one species (Whittaker 1972 ). This
leads to an exponential species-area curve because most species in a community
are common and occur within a small subsample of the area, leading to a sharp
increase in diversity between 1 and 100 m 2 evident by a linear fit to a semi-log plot
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