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200
Flagstaff
100
D
E A
0
200
Portland
100
D
S
E A
0
JFMAMJ
Month
JASOND
Fig. 10.6 Water balance for MTC site and for bimodal winter rain and summer rain
climate in Arizona. Actual evapotranspiration ( E A , dense stippling), precipitation (solid
diamonds) and potential evapotranspiration (open diamonds) contribute to water deficit (D).
S, winter surplus. (From Stephenson 1990.)
equivalent to that in the MTC of Portland, Oregon ( Fig. 10.6 ). Under summer-
rain conditions, when evapotranspiration rates are substantial, plants on severe
sites may actually be exposed to summer-drought conditions and unable to fully
utilize this summer moisture input (Vankat 1989 ). A similar phenomenon is
observed in the eastern Cape region of South Africa where MTV similar to the
Cape fynbos is distributed under a climatic regime of even rainfall throughout
the year, but due to high summer evapotranspiration it exists under essentially
a winter-rainfall climate (van Wilgen 1984 ).
On any given landscape soil moisture stress can vary markedly dependent on
microsite characteristics. Plants distributed on exposed rocky ridgelines or equator-
facing slopes with shallow soils may experience soil drought stress when other
parts of the landscape are less stressed, and plants sort out along such gradients.
Thus, landscapes comprise a mosaic of plants tolerant of varying degrees of
drought stress and with different strategies for coping with water stress (e.g.
Mooney 1989 ; Mahall et al. 2010 ). Climatic seasonality plays a key role in
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