Environmental Engineering Reference
In-Depth Information
θ
θ
a
b
θ 2
θ 1
θ
θ
1
2
φ
φ
φ
φ
φ
φ
st,2
st,1
st,2
st,1
Figure 3.8. Qualitative sketch of the dependency of the threshold
θ
on the state
variable
φ
in the case of (a) positive feedback and (b) negative feedback. The values
θ 1 and
θ 2 are also shown.
where f 1 (
0 express the state-dependent growth and decay
rates, respectively. In the presence of feedback,
φ
)
>
0and f 2 (
φ
)
<
. This depen-
dence translates into a state dependence of the switching rates of DMN, k 1 (
θ
is a function of
φ
φ
)and
k 2 (
).
As in the case with no feedback (Section 2.1), if we decrease the variance of the
driving force q while maintaining constant its mean q
φ
in the zero-variance limit, q
tends to a constant deterministic value, q
q . We can now analyze two cases of
systems with (i) positive feedback (i.e., the threshold
=
θ
decreases as
φ
increases), and
(ii) negative feedback (i.e.,
increases).
(i) In the case of positive feedback, the threshold
θ
increases as
φ
φ
[see Fig. 3.8 (a)]. In these conditions the deterministic counterpart of the stochastic
dynamics depends on the relation between
θ
is a decreasing function of
θ
(
φ
)and q . We first define the maximum
and minimum possible values of
θ
by setting in the relation
θ
(
φ
) the boundaries of
φ
φ
φ
the domain of p (
). As noted, these boundaries,
2 (minimum) and
1 (maxi-
st
,
st
,
φ/
=
φ
mum), are the steady states of the two deterministic dynamics, d
d t
f 1 , 2 (
). Thus
two values
φ st , 2 ) (maximum) are obtained [see
Fig. 3.8 (a)]. If q 1 the deterministic dynamics monotonically decrease, converg-
ing to the stable state
θ 1 = θ
(
φ st , 1 ) (minimum) and
θ 2 = θ
(
φ st , 2 . Analogously, if q 2 the system persists in state 1
(growth) regardless of the value of
φ
. Thus
φ
( t ) converges to the deterministic stable
state
q 2 . In this case the deter-
ministic system is bistable: If the threshold value associated with the initial condition
φ 0 is smaller than q
φ st , 1 . The most interesting situation is with
θ 1 <
[i.e.,
θ
(
φ 0 )
<
q ], the dynamics of
φ
exhibit a deterministic
growth with rate determined by f 1 (
) decreases and the system
persists in the growth conditions, thereby converging to the steady state,
φ
). As
φ
grows,
θ
(
φ
φ st , 1 . Con-
versely, if the system is initially in decay (or “stressed”) conditions [i.e.,
θ
(
φ 0 )
>
q ],
φ
decreases with rate f 2 (
φ
). As
φ
decreases,
θ
(
φ
) increases. Thus
φ
tends to the
steady state
φ st , 2 in that the system persists in state 2 (decay).
(ii) In the presence of negative feedback,
θ
(
φ
) increases with
φ
[see Fig. 3.8 (b)].
The two limiting values of
θ
are again defined as
θ
= θ
(
φ
1 )and
θ
= θ
(
φ
2 ).
1
st
,
2
st
,
However,
θ
1 is now the maximum and
θ
2 the minimum value of
θ
[see Fig. 3.8 (b)].
 
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