Agriculture Reference
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mass is 200-1500 kg/ha (up to 900 kg/ha in vineyards), whereas the microbial
biomass is 0.5-2 t C/ha.
SOM serves as a reserve of C and other essential elements for successive genera-
tions of organisms. Elements sequestered in the biomass are said to be immobi-
lized . Any surplus released in inorganic form is mineralized . For N, a substrate
C:N ratio 25 favors an increase in microbial biomass and net immobilization ,
whereas a C:N ratio 25 favours net mineralization.
Humus is conveniently fractionated into fulvic acid (FA), humic acid (HA), and
a nonextractable residue of humins . The FA and HA fractions are made up of
macromolecules with a “core” of substituted aromatic and aliphatic C groups. A
high density of acidic carboxyl (-COOH) and phenolic-OH groups confers a high
CEC (150-300 cmol of charge/kg), which is pH-dependent. These acidic groups
also form stable complexes ( chelates ) with metallic cations.
The rate of decomposition of litter and SOM can be expressed in terms of the
half-life of the constituents, which ranges from 1 month to 50,000 years. The
rate of decomposition is modified by clay content, moisture, temperature, and
pH.
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