Biology Reference
In-Depth Information
Table 5.2
Relative position of
neighbors to agent
x
.
l
8
l
1
l
2
l
7
x
=
l
0
l
3
l
6
l
5
l
4
Downhill:
⎛
⎞
8
i
−
1
I
p
−
1
i
I
p
−
1
j
⎝
(
⎠
f
(
x
)
=
1
−
)
l
i
i
=
1
j
=
0
⎛
⎛
⎞
p
−
1
C
m
−
1
8
⎝
I
p
−
1
0
⎝
⎠
+
(
1
−
)
·
l
0
+
i
(
k
−
I
i
)
m
=
1
k
=
0
j
=
⎛
8
i
−
1
⎝
p
−
1
p
−
1
·
1
(
1
−
(
m
−
I
i
)
)
l
i
0
(
m
−
I
j
)
i
=
j
=
⎞
⎞
p
−
1
⎠
⎠
,
+
(
1
−
(
m
−
I
0
)
)
l
0
(5.8)
where the concentration ranges from 0 to
C
C
represents the highest possible con-
centration,
I
i
is the concentration level at neighboring patch
i
, and
l
i
is the relative
location of patch
i
from the current patch, see Table
5.2
.
I
0
is the concentration of
the current patch,
I
1
,...,
,
I
8
are the concentrations of its eight neighbors.
Next, we show that Eq. (
5.7
) describes the uphill movement.
1
⎛
⎛
⎞
1
8
8
i
−
1
p
−
⎝
⎝
(
p
−
1
p
−
1
⎠
f
(
x
)
=
−
0
(
C
−
I
i
)
1
−
(
C
−
I
i
)
)
l
i
0
(
C
−
I
j
)
i
=
i
=
1
j
=
1
1
1
1
l
0
C
−
1
8
p
−
p
−
+
−
(
C
−
I
0
)
+
−
0
(
m
−
I
i
)
m
=
1
i
=
C
8
8
p
−
1
p
−
1
×
0
(
k
−
I
i
)
1
(
1
−
(
m
−
I
i
)
)
l
i
k
=
m
+
1
i
=
i
=
⎞
⎠
.
i
−
1
p
−
1
p
−
1
×
0
(
m
−
I
j
)
+
(
1
−
(
m
−
I
0
)
)
l
0
(5.9)
j
=
It is straightforward to see that Eq. (
5.9
) describes the movement to the neighboring
patch with the highest concentration level: 1
−
i
=
0
(
p
−
1
is 0 unless at least
C
−
I
i
)
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