Legs: Legs are essentially as in male. Coxal spurs are quite similar to those of
male except that spur of IV is somewhat smaller than that of III and those of II and
III are sub-equal in length and width.
Nymph ( Figures 2.94 and 2.95 ) 55
Length measures approximately 1.0 mm, and width is around 0.65 mm.
Capitulum: Basis capitulum is approximately twice as wide as long; cornua are
moderately wide and tapered. Palpi are with outline quite similar to that of female
except for slightly greater ratio of width to length. Segment III is with ventral spur
and dorsal margins similar to those of female. Hypostome is with 2/2 dental for-
mula in files of approximately seven denticles; a moderate corona is present
Scutum: Scutum is slightly broader than long; cervical grooves are as in female;
setae number four in each scapular area, two anteriorly in central field.
Legs: Coxa are with spurs essentially as in female except that the spur of IV is
smaller and directed medially.
Larva ( Figures 2.94 and 2.95 ) 55
Larva measures approximately 0.6 mm in length and 0.4 mm in width.
Capitulum: Basis capitulum is similar to that of nymph; cornua are not apparent
in unmounted specimens but suggested in mounted specimens. Palpi are with out-
line similar to that of female and nymph except that inner margin dorsally is essen-
tially straight and the contiguous margins of segments II and III are equal in
length. Hypostome is with 2/2 dental formula in files of five denticles.
Scutum: Scutum is 1.5 times as wide as long; outline is as illustrated, with two
pairs of setae and three pairs of sensilli.
Legs: Coxa I is with very small, triangular inner spur on basal margin; II and III
each has an indefinite ridge-like spur on basal margin.
Related Species 55
This species has some features in common with H. howletti Warburton, but it is
easily distinguished because of virtual absence of palpal basolateral salience
(marked in howletti); dental formula which is 4/4 (5/5 to 6/6 in howletti); different
size and distribution of denticles; scutal outline of each sex which is shorter and
wider and punctations which are less dense than in howletti; and coxal spurs which
are somewhat larger than in howletti. The difference in length of the contiguous
margins of palpal segments II and III dorsally is absent in howletti. From
H. campanulata, this species is separated by lesser palpal width basally, longer
ventral spur of palpal segment III, straight (not curved) basal margin of male palpal
segment II, difference in length of contiguous margins of palpal segments II and
III, smaller porose areas, more elongate tarsus IV lacking dorsal hump, larger coxal
spurs, more elongate extension of spiracular plate (male), shorter and broader scu-
tum, etc. This species differs from H. bispinosa and all others with a dorsobasal