and III are small, triangular, sharply pointed; spur of coxa IV is very long, dagger-like,
directed posteriorly, and extending up to or beyond the level of anal opening.
Female ( Figures 2.56 and 2.57 ) 47,104,127
The female is 2.15
1.80 mm wide.
Capitulum: Basis capitulum is dorsally almost twice as wide as long; margins
are as in male; cornua are slightly shorter than that of male, one-third as long as
the base of basis capitulum; porose areas are oval, wide apart by almost twice
their largest diameter. Palpi are as in male except segment III dorsally with a well-
developed, triangular, posteriorly directed spur at its posterior margin. Hypostome
dental formula is 4/4, with 11
3.00 mm long, 1.35
13 denticles per file.
Scutum: Scutum is sub-circular in outline. Cervical grooves are deep, narrow,
sub-parallel, and extending up to mid-length of scutum. Punctations are moderate
in size, deep, and uniformly distributed. Marginal grooves are long, narrow, and
enclosing first two festoons. Festoons number 11. Genital area is circular in outline;
operculum is broadly U-shaped.
Legs: Legs are slightly narrower than in males. Coxa I is with a long, bluntly
pointed spur; II to IV are with small triangular, bluntly pointed spurs.
Nymph( Figures 2.56 and 2.57 ) 47
Capitulum: Basis capitulum is approximately 2 times as wide as long. Cornua are
triangular, approximately one-third as long as basis capitulum. Posterior margin
between cornua is straight. Hypostome is clavate, extends slightly beyond the level
of palpi, approximately 2.5 times as long as wide; dental formula is 2/2, sometimes
3/3 apically, with eight to ten denticles per file.
Scutum: Cervical grooves are deep anteriorly, parallel up to mid-length of scu-
tum, then divergent and shallow posteriorly. Punctations are approximately 12 each
with a minute seta in addition to six pairs of sensilli. Marginal grooves are long,
deep, and enclosing first two festoons. Festoons number 11, each bearing a seta
dorsally and ventrally except the median.
Legs: Legs are moderately long and stout. Coxa I is with a distinct triangular
pointed spur; II and III are each with ridge-like spur.
Larva ( Figures 2.56 and 2.57 ) 47
Capitulum: Basis capitulum is dorsally approximately twice as broad as long; cor-
nua are one-fourth as long as basis capituli. Each palpus is approximately 1.3 times
as long as wide, conical in outline. Hypostome is with seven to eight denticles per
Scutum: Scutum is wider than long, outline trapezoid, with moderate anterior
emargination, and posterior margin broadly rounded; cervical grooves are sub-
parallel, reaching mid-length of scutum. Punctations are six in number, each with a
single seta, in addition to three pairs of sensilli. Festoons number 11.